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Yes, if you receive a toll charge while using the rental vehicle, you will need to pay the toll plus any additional processing or administration fees charged by the rental company. They will normally deduct this straight from the credit card that you used to pay for your rental.
Some toll roads are paid in cash or by cards at staffed or automatic booths on a toll plaza, others are charged electronically via transponder or number plate to avoid queues, and many offer both payment methods. We recommend that you ask at the rental counter about the possibility of an e-collection transponder, if they are used in the area you will be driving, to make your trip smoother and reduce time spent in lines waiting to pay tolls.